ΠΠ»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ Π³Π»ΡΠΊΠΎΠ·ΠΎ-6-ΡΠΎΡΡΠ°Ρ ΠΈΠ·ΠΎΠΌΠ΅ΡΠ°Π·Ρ Ρ ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ (Polychaeta: spionidae) ΠΈ Π²ΠΎΠ·ΠΌΠΎΠΆΠ½ΡΠ΅ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΡ ΠΈΡ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ
ΠΠΏΡΠΎΠ±Π°ΡΠΈΡ ΡΠ°Π±ΠΎΡΡ. ΠΡΠ½ΠΎΠ²Π½ΡΠ΅ ΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ΠΈΡ ΡΠ°Π±ΠΎΡΡ Π΄ΠΎΠΊΠ»Π°Π΄ΡΠ²Π°Π»ΠΈΡΡ Π½Π° IV Π Π΅Π³ΠΈΠΎΠ½Π°Π»ΡΠ½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ ΠΏΠΎ Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΠΌ ΠΏΡΠΎΠ±Π»Π΅ΠΌΠ°ΠΌ ΠΌΠΎΡΡΠΊΠΎΠΉ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ ΠΈ ΡΠΊΠΎΠ»ΠΎΠ³ΠΈΠΈ (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, 2001), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ: «Evolution, Genetics, Ecology and Biodiversity» (MAPEEG) (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, ΠΠΠ‘ «ΠΠΎΡΡΠΎΠΊ», 2001), VII ΠΠ°Π»ΡΠ½Π΅Π²ΠΎΡΡΠΎΡΠ½ΠΎΠΉ ΠΌΠΎΠ»ΠΎΠ΄Π΅ΠΆΠ½ΠΎΠΉ ΡΠΊΠΎΠ»Π΅ — ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ ΠΏΠΎ Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΠΌ ΠΏΡΠΎΠ±Π»Π΅ΠΌΠ°ΠΌ Ρ ΠΈΠΌΠΈΠΈ ΠΈ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, ΠΠΠ‘ Π’ΠΠΠΠ₯… Π§ΠΈΡΠ°ΡΡ Π΅ΡΡ >
- Π‘ΠΎΠ΄Π΅ΡΠΆΠ°Π½ΠΈΠ΅
- ΠΡΠ΄Π΅ΡΠΆΠΊΠ°
- ΠΠΈΡΠ΅ΡΠ°ΡΡΡΠ°
- ΠΡΡΠ³ΠΈΠ΅ ΡΠ°Π±ΠΎΡΡ
- ΠΠΎΠΌΠΎΡΡ Π² Π½Π°ΠΏΠΈΡΠ°Π½ΠΈΠΈ
Π‘ΠΎΠ΄Π΅ΡΠΆΠ°Π½ΠΈΠ΅
- 1. ΠΠΠΠΠΠΠΠ
- 2. ΠΠΠΠΠ ΠΠΠ’ΠΠ ΠΠ’Π£Π Π«
- 2. 1. ΠΠ±ΡΠ΅ΠΏΡΠΈΠ½ΡΡΠ°Ρ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ²
- 2. 1. 1. ΠΠ»Π»Π΅Π»ΡΠ½ΡΠ΅ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ
- 2. 1. 2. ΠΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ, ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΠΌΡΠ΅ ΡΠ°Π·Π½ΡΠΌΠΈ Π³Π΅Π½Π°ΠΌΠΈ
- 2. 1. 3. ΠΠΈΠ±ΡΠΈΠ΄Π½ΡΠ΅ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ
- 2. 1. ΠΠ±ΡΠ΅ΠΏΡΠΈΠ½ΡΡΠ°Ρ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ²
- 2. 2. ΠΠΎΠ²ΡΠ΅ ΠΌΠ΅Ρ
Π°Π½ΠΈΠ·ΠΌΡ Π³Π΅Π½Π΅ΡΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΌΠ½ΠΎΠΆΠ΅ΡΡΠ²Π΅Π½Π½ΠΎΡΡΠΈ Π±Π΅Π»ΠΊΠΎΠ²ΡΡ
ΠΈΠ·ΠΎ-ΡΠΎΡΠΌ
- 2. 2. 1. ΠΠ»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΠΉ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³
- 2. 2. 2. ΠΡΠΏΠΎΠ»ΡΠ·ΠΎΠ²Π°Π½ΠΈΠ΅ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΡ ΡΠ°ΠΉΡΠΎΠ² ΠΈΠ½ΠΈΡΠΈΠ°ΡΠΈΠΈ ΡΡΠ°Π½ΡΠΊΡΠΈΠΏΡΠΈΠΈ ΠΈ ΠΏΠΎΠ»ΠΈΠ°Π΄Π΅Π½ΠΈΠ»ΠΈΡΠΎΠ²Π°Π½ΠΈΡ
- 2. 2. 3. Π Π΅Π΄Π°ΠΊΡΠΈΡΠΎΠ²Π°Π½ΠΈΠ΅ ΠΌΠ ΠΠ
- 2. 2. 4. ΠΠ»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½Π°Ρ ΠΈΠ½ΠΈΡΠΈΠ°ΡΠΈΡ ΡΡΠ°Π½ΡΠ»ΡΡΠΈΠΈ
- 2. 3. ΠΠ»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΠΉ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ Π³Π΅Π½ΠΎΠ²
- 4. 1. ΠΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ ΠΠ€Π ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄
- 4. 1. 1. Polydora brevipalpa
- 4. 1. 2. Polydora calcarea
- 4. 1. 3. Polydora manchenko
- 4. 1. 4. Polydora limicola
- 4. 1. 5. Polydora glycymerica
- 4. 1. 6. Polydora cornuta
- 4. 1. 7. Dipolydora commensalis
- 4. 1. 8. Dipolydora bidentata
- 4. 1. 9. Dipolydora carunculata. Ρ 4.1.10 Dipolydora cardalia
- 4. 2. ΠΠΎΠ»Ρ ΠΈΠ΄Π΅Π½ΡΠΈΡΠ½ΡΡ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΡ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΠ€Π-1 ΠΈ ΠΠ€Π
- 5. 1. Π’ΡΡΠ΄Π½ΠΎΡΡΠΈ ΠΈΠ½ΡΠ΅ΡΠΏΡΠ΅ΡΠ°ΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ
ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΠ€Π ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ Ρ ΠΏΠΎΠ·ΠΈΡΠΈΠΉ ΠΎΠ±ΡΠ΅ΠΏΡΠΈΠ½ΡΡΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ²
- 5. 1. 1. Π₯Π°ΡΠ°ΠΊΡΠ΅Ρ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΠΎΠΉ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€ΠΠΈ ΠΠ€Π
- 5. 1. 2. Π‘ΠΎΠΏΡΡΠΆΠ΅Π½Π½ΠΎΡΡΡ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ° ΠΠ€Π-2 Ρ ΠΏΠΎΠ»ΠΎΠΌ ΠΈ ΡΠΊΠ°Π½Π΅ΠΏΠ΅ΡΠΈΡΠΈΡΠ½ΠΎΡΡΡ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π-1 ΠΈ ΠΠ€Π
- 5. 1. 3. ΠΠΎΠ»ΡΡΠΎΠ΅ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΎΠ΅ ΡΠ°Π·Π»ΠΈΡΠΈΠ΅ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π-1 ΠΈ ΠΠ€Π
- 5. 1. 4. Π€ΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΠ΅ ΠΌΠ΅ΠΆΠ»ΠΎΠΊΡΡΠ½ΡΡ Π³ΠΈΠ±ΡΠΈΠ΄ΠΎΠ² ΠΠ€Π-1/ΠΠ€Π
- 5. 2. ΠΠ»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΠΉ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³ — ΠΌΠ΅Ρ
Π°Π½ΠΈΠ·ΠΌ, ΡΠΏΠΎΡΠΎΠ±Π½ΡΠΉ Π³Π΅Π½Π΅ΡΠΈΡΠΎΠ²Π°ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ ΠΠ€Π, Π½Π°Π±Π»ΡΠ΄Π°Π΅ΠΌΡΠ΅ Ρ ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄
- 5. 2. 1. Π‘ΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½ΠΎΡΡΡ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΠΎΠΉ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΠΈ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ²
- 5. 2. 2. ΠΠΎΠ»ΡΡΠΎΠ΅ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΎΠ΅ ΡΠ°Π·Π»ΠΈΡΠΈΠ΅ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ²
- 5. 2. 3. Π’ΠΊΠ°Π½Π΅ΡΠΏΠ΅ΡΠΈΡΠΈΡΠ½ΠΎΡΡΡ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΈ Π²ΠΎΠ·ΠΌΠΎΠΆΠ½Π°Ρ ΡΠΎΠ»Ρ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΠΎΠ³ΠΎ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³Π° Π² Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°ΡΠΈΠΈ ΠΏΠΎΠ»Π° ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄
- 5. 2. 4. ΠΠΈΠ±ΡΠΈΠ΄ΠΈΠ·Π°ΡΠΈΡ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ²
- 5. 3. ΠΠΎΠ·ΠΌΠΎΠΆΠ½ΠΎΠ΅ ΡΠ΅ΡΡΠΎΠΏΠΎΠ·ΠΎΠ½Π½ΠΎΠ΅ ΠΏΡΠΎΠΈΡΡ ΠΎΠΆΠ΄Π΅Π½ΠΈΠ΅ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ° ΠΠ€Π
ΠΠ»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ Π³Π»ΡΠΊΠΎΠ·ΠΎ-6-ΡΠΎΡΡΠ°Ρ ΠΈΠ·ΠΎΠΌΠ΅ΡΠ°Π·Ρ Ρ ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ (Polychaeta: spionidae) ΠΈ Π²ΠΎΠ·ΠΌΠΎΠΆΠ½ΡΠ΅ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΡ ΠΈΡ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ (ΡΠ΅ΡΠ΅ΡΠ°Ρ, ΠΊΡΡΡΠΎΠ²Π°Ρ, Π΄ΠΈΠΏΠ»ΠΎΠΌ, ΠΊΠΎΠ½ΡΡΠΎΠ»ΡΠ½Π°Ρ)
ΠΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ — ΠΌΠ½ΠΎΠΆΠ΅ΡΡΠ²Π΅Π½Π½ΡΠ΅ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΡΠ΅ ΡΠΎΡΠΌΡ ΠΎΠ΄Π½ΠΎΠ³ΠΎ ΠΈ ΡΠΎΠ³ΠΎ ΠΆΠ΅ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ°, ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΠ΅ΠΌΡΠ΅ Π² ΠΎΠ΄Π½ΠΎΠΌ ΠΎΡΠ³Π°Π½ΠΈΠ·ΠΌΠ΅ ΠΈ ΠΎΠ±Π»Π°Π΄Π°ΡΡΠΈΠ΅ ΡΡ ΠΎΠ΄Π½ΠΎΠΉ ΡΡΠ½ΠΊΡΠΈΠ΅ΠΉ. Π‘ΠΎΡΠ΅ΡΠ°Ρ Π³Π΅Π»Π΅Π²ΡΠΉ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅Π· ΡΠ΅ΡΠΌΠ΅Π½ΡΡΠΎΠ΄Π΅ΡΠΆΠ°ΡΠΈΡ ΠΏΡΠ΅ΠΏΠ°ΡΠ°ΡΠΎΠ² Ρ ΠΏΠΎΡΠ»Π΅Π΄ΡΡΡΠΈΠΌ ΠΎΠΊΡΠ°ΡΠΈΠ²Π°Π½ΠΈΠ΅ΠΌ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ Π³Π΅Π»Π΅ΠΉ Π½Π° ΡΠΏΠ΅ΡΠΈΡΠΈΡΠ΅ΡΠΊΡΡ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ°ΡΠΈΠ²Π½ΡΡ Π°ΠΊΡΠΈΠ²Π½ΠΎΡΡΡ ΠΌΠΎΠΆΠ½ΠΎ Π²ΡΡΠ²Π»ΡΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ Π² Π²ΠΈΠ΄Π΅ ΠΎΠΊΡΠ°ΡΠ΅Π½Π½ΡΡ Π·ΠΎΠ½. ΠΠ»Π°Π³ΠΎΠ΄Π°ΡΡ ΡΠΎΠΌΡ, ΡΡΠΎ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ ΠΏΡΠΎΡΠ²Π»ΡΡΡΡΡ ΠΊΠΎΠ΄ΠΎΠΌΠΈΠ½Π°Π½ΡΠ½ΠΎ, Ρ. Π΅. ΠΎΠ΄ΠΈΠ½ Π°Π»Π»Π΅Π»Ρ Π½Π΅ ΠΌΠ°ΡΠΊΠΈΡΡΠ΅Ρ Π΄ΡΡΠ³ΠΎΠΉ, ΠΈ ΠΎ Π³Π΅Π½ΠΎΡΠΈΠΏΠ΅ ΠΎΡΠΎΠ±ΠΈ ΠΌΡ ΠΌΠΎΠΆΠ΅ΠΌ ΡΡΠ΄ΠΈΡΡ ΠΏΠΎ Π²ΡΡΠ²Π»ΡΠ΅ΠΌΠΎΠΌΡ ΠΈΠ·ΠΎ-ΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΠΎΠΌΡ ΡΠΏΠ΅ΠΊΡΡΡ, ΠΎΠ½ΠΈ ΡΠ²Π»ΡΡΡΡΡ ΡΠ΄ΠΎΠ±Π½ΡΠΌΠΈ ΠΌΠ°ΡΠΊΠ΅ΡΠ°ΠΌΠΈ Π³Π΅Π½ΠΎΠ², ΠΈ Π΄Π°Π²Π½ΠΎ ΠΈΡΠΏΠΎΠ»ΡΠ·ΡΡΡΡΡ Π² ΡΠ°Π·Π»ΠΈΡΠ½ΡΡ ΠΎΠ±Π»Π°ΡΡΡΡ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ ΠΈ ΠΌΠ΅Π΄ΠΈΡΠΈΠ½Ρ. ΠΠ°ΠΈΠ±ΠΎΠ»Π΅Π΅ ΡΠΈΡΠΎΠΊΠΎΠ΅ ΠΏΡΠΈΠΌΠ΅Π½Π΅Π½ΠΈΠ΅ ΠΎΠ½ΠΈ Π½Π°ΡΠ»ΠΈ Π² ΠΏΠΎΠΏΡΠ»ΡΡΠΈΠΎΠ½Π½ΠΎΠΉ Π³Π΅Π½Π΅ΡΠΈΠΊΠ΅ Π΄Π»Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΠΈ ΠΏΡΠΈΡΠΎΠ΄Π½ΡΡ ΠΏΠΎΠΏΡΠ»ΡΡΠΈΠΉ ΠΈ Π²ΠΈΠ΄ΠΎΠ².
ΠΠ΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠ°Ρ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ², ΠΈΡΠΏΠΎΠ»ΡΠ·ΡΠ΅ΠΌΠ°Ρ ΠΊΠ°ΠΊ ΠΎΡΠ½ΠΎΠ²Π° Π΄Π»Ρ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΈΠ½ΡΠ΅ΡΠΏΡΠ΅ΡΠ°ΡΠΈΠΈ ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠΎΠ² ΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΠΎΠ³ΠΎ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅Π·Π°, Π±ΡΠ»Π° ΡΡΠΎΡΠΌΡΠ»ΠΈΡΠΎΠ²Π°Π½Π° Π΅ΡΠ΅ Π² 70-Π΅ Π³ΠΎΠ΄Ρ, ΠΈ Π½Π° ΠΏΡΠΎΡΡΠΆΠ΅Π½ΠΈΠΈ Π±ΠΎΠ»Π΅Π΅ ΡΡΠ΅Ρ Π΄Π΅ΡΡΡΠΈΠ»Π΅ΡΠΈΠΉ ΡΡΡΠ΅ΡΡΠ²ΡΠ΅Ρ Π² Π½Π΅ΠΈΠ·ΠΌΠ΅Π½Π½ΠΎΠΉ Π²ΠΈΠ΄Π΅ (Harris and Hopkinson, 1976; ΠΠΎΡΠΎΡΠΊΠΈΠ½ ΠΈ Π΄Ρ., 1977). ΠΡΠ° ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΡ Π²ΡΠ΄Π΅Π»ΡΠ΅Ρ ΡΡΠΈ ΠΎΡΠ½ΠΎΠ²Π½ΡΠ΅ ΠΏΡΠΈΡΠΈΠ½Ρ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈ Π²ΡΡΠ²Π»ΡΠ΅ΠΌΠΎΠΉ ΠΌΠ½ΠΎΠΆΠ΅ΡΡΠ²Π΅Π½Π½ΠΎΡΡΠΈ ΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ². ΠΠΎ-ΠΏΠ΅ΡΠ²ΡΡ , ΡΡΠΎ Π³Π΅Π½Π½ΡΠ΅ Π΄ΡΠΏΠ»ΠΈΠΊΠ°ΡΠΈΠΈ, Π²Π΅Π΄ΡΡΠΈΠ΅ ΠΊ Π²ΠΎΠ·Π½ΠΈΠΊΠ½ΠΎΠ²Π΅Π½ΠΈΡ Π½Π΅Π·Π°Π²ΠΈΡΠΈΠΌΡΡ Π³Π΅Π½Π½ΡΡ Π»ΠΎΠΊΡΡΠΎΠ², ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΡΡΠΈΡ ΡΠ°Π·Π½ΡΠ΅ ΠΈΠ·ΠΎΡΠΎΡΠΌΡ ΠΎΠ΄Π½ΠΎΠ³ΠΎ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ°, ΠΊΠΎΡΠΎΡΡΠ΅ ΠΈ ΡΠ΅ΠΊΠΎΠΌΠ΅Π½Π΄ΠΎΠ²Π°Π½ΠΎ Π½Π°Π·ΡΠ²Π°ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°ΠΌΠΈ. ΠΠΎ-Π²ΡΠΎΡΡΡ , ΡΡΠΎ Π³Π΅Π½Π½ΡΠ΅ ΠΌΡΡΠ°ΡΠΈΠΈ, ΠΏΡΠΈΠ²ΠΎΠ΄ΡΡΠΈΠ΅ ΠΊ Π²ΠΎΠ·Π½ΠΈΠΊΠ½ΠΎΠ²Π΅Π½ΠΈΡ ΠΌΠ½ΠΎΠΆΠ΅ΡΡΠ²Π΅Π½Π½ΡΡ Π²Π°ΡΠΈΠ°Π½ΡΠΎΠ² ΠΎΠ΄Π½ΠΎΠ³ΠΎ Π³Π΅Π½Π½ΠΎΠ³ΠΎ Π»ΠΎΠΊΡΡΠ°, ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΡΡΠΈΡ Π°Π»Π»Π΅Π»ΡΠ½ΡΠ΅ Π²Π°ΡΠΈΠ°Π½ΡΡ ΠΎΠ΄Π½ΠΎΠ³ΠΎ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ°, ΠΏΠΎΠ»ΡΡΠΈΠ²ΡΠΈΡ Π½Π°Π·Π²Π°Π½ΠΈΠ΅ «Π°Π»Π»ΠΎΠ·ΠΈΠΌΠΎΠ²». Π-ΡΡΠ΅ΡΡΠΈΡ , ΡΡΠΎ ΠΏΠΎΡΠΏΡΠ°Π½ΡΠ»ΡΡΠΈΠΎΠ½Π½ΡΠ΅ ΠΌΠΎΠ΄ΠΈΡΠΈΠΊΠ°ΡΠΈΠΈ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ», ΠΊΠΎΡΠΎΡΡΠ΅ ΠΏΡΠΈΠ²ΠΎΠ΄ΡΡ ΠΊ Π²ΠΎΠ·Π½ΠΈΠΊΠ½ΠΎΠ²Π΅Π½ΠΈΡ Π²ΡΠΎΡΠΈΡΠ½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ², ΠΈΠΌΠ΅ΡΡΠΈΡ Π½Π΅Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΡΡ, ΠΌΠΎΠ΄ΠΈΡΠΈΠΊΠ°ΡΠΈΠΎΠ½Π½ΡΡ ΠΏΡΠΈΡΠΎΠ΄Ρ.
Π―ΡΠ½Π°Ρ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠ°Ρ ΠΈΠ½ΡΠ΅ΡΠΏΡΠ΅ΡΠ°ΡΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΊΠ»ΡΡΠ΅Π²ΡΠΌ ΠΌΠΎΠΌΠ΅Π½ΡΠΎΠΌ Π² ΠΈΡΠΏΠΎΠ»ΡΠ·ΠΎΠ²Π°Π½ΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΊΠ°ΠΊ ΠΌΠ°ΡΠΊΠ΅ΡΠΎΠ² Π³Π΅Π½ΠΎΠ², ΠΎΠ΄Π½Π°ΠΊΠΎ ΠΎΠ½Π° Π²ΡΠ΅Π³Π΄Π° ΠΈΠΌΠ΅Π»Π° ΠΎΠΏΡΠ΅Π΄Π΅Π»Π΅Π½Π½ΡΠ΅ ΠΏΡΠΎΠ±Π»Π΅ΠΌΡ. ΠΠ½Π°ΡΠΈΡΠ΅Π»ΡΠ½Π°Ρ ΡΠ°ΡΡΡ ΡΡΠΈΡ ΠΏΡΠΎΠ±Π»Π΅ΠΌ ΠΎΡΠ½ΠΎΡΠΈΡΡΡ ΠΊ ΡΠ°Π·ΡΡΠ΄Ρ ΠΌΠ΅ΡΠΎΠ΄ΠΈΡΠ΅ΡΠΊΠΈΡ ΠΈ Π²ΡΡΠ°ΠΆΠ°Π΅ΡΡΡ Π² Π½Π΅Π΄ΠΎΡΡΠ°ΡΠΎΡΠ½ΠΎ ΡΠ΅ΡΠΊΠΎΠΌ ΡΠ°Π·Π΄Π΅Π»Π΅Π½ΠΈΠΈ Π·ΠΎΠ½ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΠΎΠΉ Π°ΠΊΡΠΈΠ²Π½ΠΎΡΡΠΈ Π½Π° ΡΠΏΠ΅ΡΠΈΡΠΈΡΠ΅ΡΠΊΠΈ ΠΎΠΊΡΠ°ΡΠ΅Π½Π½ΡΡ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ Π³Π΅Π»ΡΡ — Π·ΠΈΠΌΠΎΠ³ΡΠ°ΠΌΠΌΠ°Ρ . ΠΠ΄Π½Π°ΠΊΠΎ Π΄ΡΡΠ³Π°Ρ ΡΠ°ΡΡΡ Π²ΠΊΠ»ΡΡΠ°Π΅Ρ ΠΏΡΠΎΠ±Π»Π΅ΠΌΡ, ΡΠ²ΡΠ·Π°Π½Π½ΡΠ΅ Ρ Π½Π΅ΡΠΎΠ²Π΅ΡΡΠ΅Π½ΡΡΠ²ΠΎΠΌ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΠΈ ΠΈ Π½Π΅Π΄ΠΎΡΡΠ°ΡΠΎΡΠ½ΠΎΡΡΡΡ Π½Π°ΡΠΈΡ Π·Π½Π°Π½ΠΈΠΉ ΠΎ ΠΏΡΠΈΡΠΎΠ΄Π΅ ΠΈΠ·ΠΎΡΠ΅Ρ-ΠΌΠ΅Π½ΡΠΎΠ² ΠΈ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠ°Ρ ΠΈΡ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ. Π Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΠ΅ ΠΎΠΏΠΈΡΠ°Π½ΠΎ ΠΌΠ½ΠΎΠ³ΠΎ Π½Π΅ΠΎΠ±ΡΡΠ½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ ΡΠΏΠ΅ΠΊΡΡΠΎΠ², Π½Π΅ ΠΈΠΌΠ΅ΡΡΠΈΡ ΠΎΠ±ΠΎΡΠ½ΠΎΠ²Π°Π½Π½ΠΎΠΉ ΠΈ ΡΠ±Π΅Π΄ΠΈΡΠ΅Π»ΡΠ½ΠΎΠΉ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΈΠ½ΡΠ΅ΡΠΏΡΠ΅ΡΠ°ΡΠΈΠΈ. Π ΡΠΈΡΠ»Ρ ΡΠ°ΠΊΠΈΡ ΡΠ»ΡΡΠ°Π΅Π² ΠΎΡΠ½ΠΎΡΠΈΡΡΡ Π½Π΅Π΄Π°Π²Π½ΠΎ ΠΎΠΏΠΈΡΠ°Π½Π½ΡΠΉ Π½Π΅ΠΎΠ±ΡΡΠ½ΡΠΉ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΠΉ ΡΠΏΠ΅ΠΊΡΡ Π΄ΠΈΠΌΠ΅ΡΠ½ΠΎΠ³ΠΎ ΡΠ΅ΡΠΌΠ΅Π½ΡΠ° Π³Π»ΡΠΊΠΎΠ·ΠΎ-6-ΡΠΎΡΡΠ°Ρ ΠΈΠ·ΠΎΠΌΠ΅ΡΠ°Π·Ρ (ΠΠ€Π) Ρ ΠΏΠΎΠ»ΠΈΡ Π΅ΡΡ Polydora brevipalpa (Manchenko, 2001Π°). ΠΡΠ»ΠΎ Π²ΡΡΠ²Π»Π΅Π½ΠΎ Π΄Π²Π° ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ° ΠΠ€Π: ΠΠ€Π-1 ΠΈ ΠΠ€Π-2. ΠΠ½ΠΎΠ΄Π½ΡΠΉ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½Ρ ΠΠ€Π-1 Π²Ρ* ΡΠ²Π»ΡΠ»ΡΡ Ρ Π²ΡΠ΅Ρ ΠΎΡΠΎΠ±Π΅ΠΉ, Π° ΠΊΠ°ΡΠΎΠ΄Π½ΡΠΉ ΠΠ€Π-2 — ΡΠΎΠ»ΡΠΊΠΎ Ρ ΡΠ°ΠΌΡΠΎΠ². ΠΠ΅ΠΎΠ±ΡΡΠ½ΠΎΡΡΡ ΡΠΏΠ΅ΠΊΡΡΠ° ΡΠΎΡΡΠΎΡΠ»Π° Π² ΡΠΎΠΌ, ΡΡΠΎ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ ΠΏΡΠΎΡΠ²Π»ΡΠ»ΠΈ ΡΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½ΡΡ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΡ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡ (ΠΈΠΌΠ΅Π»ΠΈ ΠΈΠ΄Π΅Π½ΡΠΈΡΠ½ΡΠ΅ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ) ΠΈ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π»ΠΈ Π³ΠΈΠ±ΡΠΈΠ΄Ρ. Π‘ΠΏΠΎΡΠΎΠ±Π½ΠΎΡΡΡ ΠΎΠ±ΡΠ°Π·ΠΎΠ²ΡΠ²Π°ΡΡ Π³ΠΈΠ±ΡΠΈΠ΄Ρ in vivo ΠΏΡΠΈΡΡΡΠ° ΡΠΎΠ»ΡΠΊΠΎ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°ΠΌ, ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΠΌΡΠΌ Π΄ΡΠΏΠ»ΠΈΡΠΈΡΠΎΠ²Π°Π½Π½ΡΠΌΠΈ Π³Π΅Π½Π½ΡΠΌΠΈ Π»ΠΎΠΊΡΡΠ°ΠΌΠΈ, Π° ΡΠΏΠΎΡΠΎΠ±Π½ΠΎΡΡΡ ΠΏΡΠΎΡΠ²Π»ΡΡΡ ΡΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½ΡΡ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΡ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡ — ΡΠΎΠ»ΡΠΊΠΎ Π²ΡΠΎΡΠΈΡΠ½ΡΠΌ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°ΠΌ, ΠΎΠ±ΡΠ°Π·ΡΡΡΠΈΠΌΡΡ ΠΈΠ· ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΡΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π² ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠ΅ ΠΏΠΎΡΡΠ³ΡΠ°Π½ΡΠ»ΡΡΠΈΠΎΠ½Π½ΡΡ ΠΌΠΎΠ΄ΠΈΡΠΈΠΊΠ°ΡΠΈΠΉ (ΠΠΎΡΠΎΡΠΊΠΈΠ½ ΠΈ Π΄Ρ., 1977; Moss, 1982; Π€ΠΈΠ»ΠΈΠΏΠΏΠΎΠ²ΠΈΡ ΠΈ ΠΠΎΠ½ΠΈΡΠ΅Π², 1987). Π’Π°ΠΊΠΈΠΌ ΠΎΠ±ΡΠ°Π·ΠΎΠΌ, ΠΎΠ±ΡΠ΅ΠΏΡΠΈΠ½ΡΡΠ°Ρ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² Π½Π΅ ΠΌΠΎΠ³Π»Π° Π΄Π°ΡΡ ΡΠ΄ΠΎΠ²Π»Π΅ΡΠ²ΠΎΡΠΈΡΠ΅Π»ΡΠ½ΠΎΠΉ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΈΠ½ΡΠ΅ΡΠΏΡΠ΅ΡΠ°ΡΠΈΠΈ ΡΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½ΠΎΠΉ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΠΎΠΉ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΠΈ Π³ΠΈΠ±ΡΠΈΠ΄ΠΈΠ·ΡΡΡΠΈΡ ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ². ΠΠ»Ρ ΠΎΠ±ΡΡΡΠ½Π΅Π½ΠΈΡ Π½Π΅ΠΎΠ±ΡΡΠ½ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΠ€Π Polydora brevipalpa ΠΠ°Π½ΡΠ΅Π½ΠΊΠΎ (Manchenko, 2001Π°) ΠΏΡΠ΅Π΄Π»ΠΎΠΆΠΈΠ» ΠΌΠ΅f Ρ Π°Π½ΠΈΠ·ΠΌ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΠΎΠ³ΠΎ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³Π° (ΠΠ‘). ΠΠ·Π²Π΅ΡΡΠ½ΠΎ, ΡΡΠΎ ΠΠ‘ ΠΏΡΠ΅-ΠΌΠ ΠΠ ΡΡΠ°Π½Ρ-ΠΊΡΠΈΠΏΡΠΎΠ², ΡΡΠΈΡΡΠ²Π°Π΅ΠΌΡΡ Ρ ΠΎΠ΄Π½ΠΎΠ³ΠΎ ΡΡΡΡΠΊΡΡΡΠ½ΠΎΠ³ΠΎ Π³Π΅Π½Π°, ΠΌΠΎΠΆΠ΅Ρ Π΄Π°Π²Π°ΡΡ Π½Π΅ΡΠΊΠΎΠ»ΡΠΊΠΎ ΡΡΡΡΠΊΡΡΡΠ½ΠΎ ΠΎΡΠ»ΠΈΡΠ°ΡΡΠΈΡ ΡΡ Π±Π΅Π»ΠΊΠΎΠ²ΡΡ ΠΈΠ·ΠΎΡΠΎΡΠΌ (Andreadis et al., 1987). ΠΠ»Ρ ΠΎΠ±ΠΎΠ·Π½Π°ΡΠ΅Π½ΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ², Π³Π΅Π½Π΅ΡΠΈΡΡΠ΅ΠΌΡΡ ΠΠ‘, Π±ΡΠ» ΠΏΡΠ΅Π΄Π»ΠΎΠΆΠ΅Π½ Π½ΠΎΠ²ΡΠΉ ΡΠ΅ΡΠΌΠΈΠ½ «Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΠ΅ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ» (Manchenko, 2001Π°). Π‘ΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½Π°Ρ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½Π°Ρ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΈ ΠΈΡ ΡΠΏΠΎΡΠΎΠ±Π½ΠΎΡΡΡ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°ΡΡ Π³ΠΈΠ±ΡΠΈΠ΄Ρ Π±ΡΠ»ΠΈ ΡΠ°ΡΡΠ΅Π½Π΅Π½Ρ ΠΊΠ°ΠΊ Π²Π΅ΡΠΊΠΈΠ΅ Π°ΡΠ³ΡΠΌΠ΅Π½ΡΡ Π² ΠΏΠΎΠ»ΡΠ·Ρ ΡΡΠ°ΡΡΠΈΡ ΠΠ‘ Π² Π³Π΅Π½Π΅ΡΠΈΡΠΎΠ²Π°Π½ΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π P. brevipalpa (Manchenko, 2001Π°). ΠΠ΄Π½Π°ΠΊΠΎ ΠΏΡΠ΅Π΄Π²Π°ΡΠΈΡΠ΅Π»ΡΠ½ΡΠ΅ Π΄Π°Π½Π½ΡΠ΅ ΡΠ²ΠΈΠ΄Π΅ΡΠ΅Π»ΡΡΡΠ²ΠΎΠ²Π°Π»ΠΈ ΠΎ ΡΠΎΠΌ, ΡΡΠΎ Ρ Π±Π»ΠΈΠ·ΠΊΠΈΡ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ P. ciliata sp. l ΠΈ P. limicola * ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ ΠΠ€Π ΠΏΡΠΎΡΠ²Π»ΡΡΡ Π½Π΅ΡΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½ΡΡ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΡ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡ, Ρ Π°ΡΠ°ΠΊΡΠ΅ΡΠ½ΡΡ Π΄Π»Ρ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ², ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΠΌΡΡ Π΄Π²ΡΠΌΡ ΡΠ°Π·Π½ΡΠΌΠΈ Π³Π΅Π½Π°ΠΌΠΈ (Manchenko and Radashevsky, 1993,1998).
Π¦Π΅Π»Ρ Π½Π°ΡΡΠΎΡΡΠ΅ΠΉ ΡΠ°Π±ΠΎΡΡ Π·Π°ΠΊΠ»ΡΡΠ°Π»Π°ΡΡ Π² Π²ΡΡΡΠ½Π΅Π½ΠΈΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΡΡ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠΎΠ² ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΠ€Π Ρ ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ (Polychaeta: Spi-onidae).
ΠΠ»Ρ Π΄ΠΎΡΡΠΈΠΆΠ΅Π½ΠΈΡ ΠΏΠΎΡΡΠ°Π²Π»Π΅Π½Π½ΠΎΠΉ ΡΠ΅Π»ΠΈ Π±ΡΠ»ΠΈ ΠΎΠΏΡΠ΅Π΄Π΅Π»Π΅Π½Ρ ΡΠ»Π΅Π΄ΡΡΡΠΈΠ΅ Π·Π°Π΄Π°ΡΠΈ:
1) ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ ΠΠ€Π Ρ Π΄Π΅ΡΡΡΠΈ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ (Polychaeta: Spionidae).
2) ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΡ ΡΠΎΠΏΡΡΠΆΠ΅Π½Π½ΠΎΡΡΡ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ° ΠΠ€Π-2 Ρ ΠΏΠΎΠ»ΠΎΠΌ Ρ Π΄Π΅ΡΡΡΠΈ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄.
3) ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΡ ΡΠΊΠ°Π½Π΅ΡΠΏΠ΅ΡΠΈΡΠΈΡΠ½ΠΎΡΡΡ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π-1 ΠΈ ΠΠ€Π-2.
ΠΠ°ΡΡΠ½Π°Ρ Π½ΠΎΠ²ΠΈΠ·Π½Π° ΡΠ°Π±ΠΎΡΡ. ΠΡΠΎΠ²Π΅Π΄Π΅Π½ΠΎ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΎΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ½ΡΡ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² Π³Π»ΡΠΊΠΎΠ·ΠΎ-6-ΡΠΎΡΡΠ°Ρ ΠΈΠ·ΠΎΠΌΠ΅ΡΠ°Π·Ρ (ΠΠ€Π) Ρ Π΄Π΅ΡΡΡΠΈ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ ΡΠΎΠ΄ΠΎΠ² Polydora ΠΈ Dipolydora (Polychaeta: Spionidae), ΡΠ°Π½Π΅Π΅ Π½Π΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π² ΡΡΠΎΠΌ ΠΎΡΠ½ΠΎΡΠ΅Π½ΠΈΠΈ. Π£ΡΡΠ°Π½ΠΎΠ²Π»Π΅Π½ΠΎ, ΡΡΠΎ Ρ Π²ΡΠ΅Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΠ΅ΡΡΡ Π΄Π²Π° ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ° (ΠΠ€Π-1 ΠΈ ΠΠ€Π-2), ΠΎΠ΄ΠΈΠ½ ΠΈΠ· ΠΊΠΎΡΠΎΡΡΡ (ΠΠ€Π-2) ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΠ΅ΡΡΡ ΠΏΠΎΡΡΠΈ ΠΈΡΠΊΠ»ΡΡΠΈΡΠ΅Π»ΡΠ½ΠΎ Ρ ΡΠ°ΠΌΡΠΎΠ². ΠΡΠΎΠ²Π΅Π΄Π΅Π½ΠΎ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠΊΠ°Π½Π΅ΡΠΏΠ΅ΡΠΈΡΠΈΡΠ½ΠΎΡΡΠΈ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π. ΠΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ ΠΈΠ·ΠΎΡΠ΅Ρ-ΠΌΠ΅Π½Ρ ΠΠ€Π-1 ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΠ΅ΡΡΡ ΠΏΡΠ΅ΠΈΠΌΡΡΠ΅ΡΡΠ²Π΅Π½Π½ΠΎ Π² ΡΠΎΠΌΠ°ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠΊΠ°Π½ΡΡ Ρ ΠΎΡΠΎΠ±Π΅ΠΉ ΠΎΠ±ΠΎΠΈΡ ΠΏΠΎΠ»ΠΎΠ² ΠΈ Π² Π½Π΅Π·Π½Π°ΡΠΈΡΠ΅Π»ΡΠ½ΡΡ ΠΊΠΎΠ»ΠΈΡΠ΅ΡΡΠ²Π°Ρ Π² Π΄ΠΈΠΏΠ»ΠΎΠΈΠ΄Π½ΡΡ ΡΠΏΠ΅ΡΠΌΠ°ΡΠΎΠ³Π΅Π½Π½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΡΠ°ΠΌΡΠΎΠ². ΠΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½Ρ ΠΠ€Π-2 ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΠ΅ΡΡΡ Π² Π΄ΠΈΠΏΠ»ΠΎΠΈΠ΄Π½ΡΡ ΡΠΏΠ΅ΡΠΌΠ°ΡΠΎΠ³Π΅Π½Π½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΡΠ°ΠΌΡΠΎΠ² ΠΈ Π² ΡΠ»Π΅Π΄ΠΎΠ²ΡΡ ΠΊΠΎΠ»ΠΈΡΠ΅ΡΡΠ²Π°Ρ Ρ ΡΠ°ΠΌΠΎΠΊ. Π£ΡΡΠ°Π½ΠΎΠ²Π»Π΅Π½ ΡΠ°ΠΊΡ Π³ΠΈΠ±ΡΠΈΠ΄ΠΈΠ·Π°ΡΠΈΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π-1 ΠΈ ΠΠ€Π-2 Ρ Π²ΡΠ΅Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ². ΠΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ Ρ ΠΊΠ°ΠΆΠ΄ΠΎΠ³ΠΎ ΠΈΠ· ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ ΡΡΠ΅Π΄ΠΈ ΠΎΡΠΎΠ±Π΅ΠΉ, ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΡ-ΡΠΈΡ ΠΎΠ±Π° ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°, ΠΏΡΠ΅ΠΎΠ±Π»Π°Π΄Π°ΡΡ ΡΠ΅, ΠΊΠΎΡΠΎΡΡΠ΅ ΠΈΠΌΠ΅ΡΡ ΠΈΠ΄Π΅Π½ΡΠΈΡΠ½ΡΠ΅ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΡΠ΅ ΡΠΏΠ΅ΠΊΡΡΡ ΠΠ€Π-1 ΠΈ ΠΠ€Π-2. ΠΠΏΠ΅ΡΠ²ΡΠ΅ ΡΠ΄Π΅Π»Π°Π½Π° ΠΎΡΠ΅Π½ΠΊΠ° Π΄ΠΎΠ»ΠΈ ΡΠ°ΠΊΠΈΡ ΠΎΡΠΎΠ±Π΅ΠΉ Ρ 10 ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ².
ΠΠΏΠ΅ΡΠ²ΡΠ΅ ΠΏΠΎΠ»ΡΡΠ΅Π½Ρ Π΄Π°Π½Π½ΡΠ΅, ΡΠ²ΠΈΠ΄Π΅ΡΠ΅Π»ΡΡΡΠ²ΡΡΡΠΈΠ΅ Π² ΠΏΠΎΠ»ΡΠ·Ρ Π³ΠΈΠΏΠΎΡΠ΅Π·Ρ ΠΎ ΡΠΎΠΌ, ΡΡΠΎ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΡΠΉ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³ ΡΠΏΠΎΡΠΎΠ±Π΅Π½ Π³Π΅Π½Π΅ΡΠΈΡΠΎΠ²Π°ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°, ΠΊΠΎΡΠΎΡΡΠ΅ Π½Π° ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ Π³Π΅Π»ΡΡ ΠΌΠΎΠ³ΡΡ Π±ΡΡΡ Π½Π΅ΠΎΡΠ»ΠΈΡΠΈΠΌΡ ΠΎΡ ΠΊΠ»Π°ΡΡΠΈΡΠ΅ΡΠΊΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ², ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΠΌΡΡ ΡΠ°Π·Π½ΡΠΌΠΈ Π³Π΅Π½Π½ΡΠΌΠΈ Π»ΠΎΠΊΡΡΠ°ΠΌΠΈ.
Π’Π΅ΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠ°Ρ ΠΈ ΠΏΡΠ°ΠΊΡΠΈΡΠ΅ΡΠΊΠ°Ρ ΡΠ΅Π½Π½ΠΎΡΡΡ ΡΠ°Π±ΠΎΡΡ. ΠΠΎΠ»ΡΡΠ΅Π½Π½ΡΠ΅ Π΄Π°Π½Π½ΡΠ΅, Π³ΠΎΠ²ΠΎΡΡΡΠΈΠ΅ ΠΎ ΡΠΎΠΌ, ΡΡΠΎ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ ΠΠ€Π ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ Π³Π΅Π½Π΅ΡΠΈΡΡΡΡΡΡ ΠΠ‘, ΠΏΠΎΡΠ»ΡΠΆΠ°Ρ Π²Π°ΠΆΠ½ΠΎΠΉ ΠΏΡΠ΅Π΄ΠΏΠΎΡΡΠ»ΠΊΠΎΠΉ Π΄Π»Ρ ΡΠΎΡΠΌΡΠ»ΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ΠΈΠΉ Π½ΠΎΠ²ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΠΈ ΠΈΠ·ΠΎ-ΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ², ΡΡΠΈΡΡΠ²Π°ΡΡΠ΅ΠΉ Π²Π°ΠΆΠ½ΡΡ ΡΠΎΠ»Ρ ΠΠ‘ Π² ΠΈΡ Π³Π΅Π½Π΅ΡΠΈΡΠΎΠ²Π°Π½ΠΈΠΈ. ΠΡΠΎ ΡΠΎΡΡΠ°Π²ΠΈΡ ΠΎΡΠ½ΠΎΠ²Ρ Π΄Π»Ρ ΡΡΠ½ΠΎΠΉ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΈΠ½ΡΠ΅ΡΠΏΡΠ΅ΡΠ°ΡΠΈΠΈ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ Π΄Π°Π½Π½ΡΡ ΠΈ ΠΏΠΎΠ΄ΡΠ²Π΅ΡΠ΄ΠΈΡ Π΄Π»Ρ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΡΡΠ°ΡΡΡ Π½Π°Π΄Π΅ΠΆΠ½ΡΡ Π³Π΅Π½Π½ΡΡ ΠΌΠ°ΡΠΊΠ΅ΡΠΎΠ².
ΠΠΏΡΠΎΠ±Π°ΡΠΈΡ ΡΠ°Π±ΠΎΡΡ. ΠΡΠ½ΠΎΠ²Π½ΡΠ΅ ΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ΠΈΡ ΡΠ°Π±ΠΎΡΡ Π΄ΠΎΠΊΠ»Π°Π΄ΡΠ²Π°Π»ΠΈΡΡ Π½Π° IV Π Π΅Π³ΠΈΠΎΠ½Π°Π»ΡΠ½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ ΠΏΠΎ Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΠΌ ΠΏΡΠΎΠ±Π»Π΅ΠΌΠ°ΠΌ ΠΌΠΎΡΡΠΊΠΎΠΉ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ ΠΈ ΡΠΊΠΎΠ»ΠΎΠ³ΠΈΠΈ (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, 2001), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ: «Evolution, Genetics, Ecology and Biodiversity» (MAPEEG) (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, ΠΠΠ‘ «ΠΠΎΡΡΠΎΠΊ», 2001), VII ΠΠ°Π»ΡΠ½Π΅Π²ΠΎΡΡΠΎΡΠ½ΠΎΠΉ ΠΌΠΎΠ»ΠΎΠ΄Π΅ΠΆΠ½ΠΎΠΉ ΡΠΊΠΎΠ»Π΅ — ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ ΠΏΠΎ Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΠΌ ΠΏΡΠΎΠ±Π»Π΅ΠΌΠ°ΠΌ Ρ ΠΈΠΌΠΈΠΈ ΠΈ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, ΠΠΠ‘ Π’ΠΠΠΠ₯, 2003) ΠΈ Π΅ΠΆΠ΅Π³ΠΎΠ΄Π½ΡΡ Π½Π°ΡΡΠ½ΡΡ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΡΡ ΠΠ½ΡΡΠΈΡΡΡΠ° Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ ΠΌΠΎΡΡ (ΠΠ»Π°Π΄ΠΈΠ²ΠΎΡΡΠΎΠΊ, 2004,2006).
Π‘ΡΡΡΠΊΡΡΡΠ° ΠΈ ΠΎΠ±ΡΠ΅ΠΌ ΡΠ°Π±ΠΎΡΡ. Π Π°Π±ΠΎΡΠ° ΡΠΎΡΡΠΎΠΈΡ ΠΈΠ· Π²Π²Π΅Π΄Π΅Π½ΠΈΡ, ΠΎΠ±Π·ΠΎΡΠ° Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΡ, ΠΎΠΏΠΈΡΠ°Π½ΠΈΡ ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π»ΠΎΠ² ΠΈ ΠΌΠ΅ΡΠΎΠ΄ΠΎΠ², ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠΎΠ² ΠΈ ΠΎΠ±ΡΡΠΆΠ΄Π΅Π½ΠΈΡ, Π²ΡΠ²ΠΎΠ΄ΠΎΠ² ΠΈ ΡΠΏΠΈΡΠΊΠ° ΡΠΈΡΠΈΡΡΠ΅ΠΌΠΎΠΉ Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΡ, ΡΠΎΠ΄Π΅ΡΠΆΠ°ΡΠ΅Π³ΠΎ 209 ΡΡΡΠ»ΠΎΠΊ. ΠΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΡ ΠΈΠ·Π»ΠΎΠΆΠ΅Π½Π° Π½Π° 108 ΡΡΡΠ°Π½ΠΈΡΠ°Ρ , ΡΠΎΠ΄Π΅ΡΠΆΠΈΡ 4 ΡΠ°Π±Π»ΠΈΡΡ ΠΈ 43 ΡΠΈΡΡΠ½ΠΊΠ°.
ΠΠ«ΠΠΠΠ«
1. Π£ Π²ΡΠ΅Ρ Π΄Π΅ΡΡΡΠΈ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ (Polydora brevipalpa, P. cal-carea, P. cornuta, P. glycymerica, P. limicola, P. manchenkoi, Dipolydora biden-tata, D. cardalia, D. carunculata, D. commensalis (Polychaeta: Spionidae)) Ρ ΡΠ°ΠΌΡΠΎΠ² Π²ΡΡΠ²Π»Π΅Π½ΠΎ Π΄Π²Π° ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ° ΠΠ€Π — ΠΠ€Π-1 ΠΈ ΠΠ€Π-2, Ρ ΡΠ°ΠΌΠΎΠΊ-ΡΠΎΠ»ΡΠΊΠΎ ΠΎΠ΄ΠΈΠ½ ΠΠ€Π-1.
2. ΠΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½Ρ ΠΠ€Π-1 Ρ Π²ΡΠ΅Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ² ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΡΡΠ΅ΡΡΡ Π² ΡΠΎΠΌΠ°ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠΊΠ°Π½ΡΡ , Π° ΠΠ€Π-2 — Π² ΡΠΏΠ΅ΡΠΌΠ΅.
3. Π£ Π²ΡΠ΅Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ Π²ΡΡΠ²Π»Π΅Π½Ρ Π³ΠΈΠ±ΡΠΈΠ΄Π½ΡΠ΅ Π·ΠΎΠ½Ρ ΠΌΠ΅ΠΆΠ΄Ρ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°ΠΌΠΈ ΠΠ€Π-1 ΠΈ ΠΠ€Π-2, ΡΡΠΎ ΡΠ²ΠΈΠ΄Π΅ΡΠ΅Π»ΡΡΡΠ²ΡΠ΅Ρ ΠΎ ΡΠΎΠΌ, ΡΡΠΎ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½Ρ ΠΠ€Π-2 Π½Π΅ ΠΌΠΎΠΆΠ΅Ρ ΠΈΠΌΠ΅ΡΡ ΠΏΠΎΡΡΡΡΠ°Π½ΡΠ»ΡΡΠΈΠΎΠ½Π½ΠΎΠ΅ ΠΏΡΠΎΠΈΡΡ ΠΎΠΆΠ΄Π΅Π½ΠΈΠ΅. ΠΠ± ΡΡΠΎΠΌ ΠΆΠ΅ ΡΠ²ΠΈΠ΄Π΅ΡΠ΅Π»ΡΡΡΠ²ΡΠ΅Ρ ΠΈ Π±ΠΎΠ»ΡΡΠΎΠ΅ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΎΠ΅ ΡΠ°Π·Π»ΠΈΡΠΈΠ΅ ΠΌΠ΅ΠΆΠ΄Ρ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠ°ΠΌΠΈ ΠΠ€Π-1 ΠΈ ΠΠ€Π-2.
4. Π‘ΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½Π°Ρ ΠΏΠΎΠ»ΠΈΠ°Π»Π»Π΅Π»ΡΠ½Π°Ρ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½Π°Ρ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² ΠΠ€Π-1 ΠΈ ΠΠ€Π-2 Ρ Π±ΠΎΠ»ΡΡΠΈΠ½ΡΡΠ²Π° ΠΎΡΠΎΠ±Π΅ΠΉ Ρ Π²ΡΠ΅Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ Π²ΠΈΠ΄ΠΎΠ² ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ ΡΠ²ΠΈΠ΄Π΅ΡΠ΅Π»ΡΡΡΠ²ΡΡΡ ΠΏΡΠΎΡΠΈΠ² ΡΠΎΠ³ΠΎ, ΡΡΠΎ ΡΡΠΈ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΡ ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΡΡΡΡ ΡΠ°Π·Π½ΡΠΌΠΈ Π³Π΅Π½Π½ΡΠΌΠΈ Π»ΠΎΠΊΡΡΠ°ΠΌΠΈ.
5. Π‘Π²ΠΎΠ΅ΠΎΠ±ΡΠ°Π·ΠΈΠ΅ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΠ€Π ΠΏΠΎΠ»ΠΈΠ΄ΠΎΡΠΈΠ΄ — ΡΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½Π°Ρ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½Π°Ρ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡ Π΄Π²ΡΡ Π³ΠΈΠ±ΡΠΈΠ΄ΠΈΠ·ΡΡΡΠΈΡ ΡΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² Ρ Π±ΠΎΠ»ΡΡΠΈΠ½ΡΡΠ²Π° ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½Π½ΡΡ ΠΎΡΠΎΠ±Π΅ΠΉ Π½Π°ΡΡΠ΄Ρ Ρ Π½Π°Π»ΠΈΡΠΈΠ΅ΠΌ ΡΠ΅Π΄ΠΊΠΈΡ ΠΎΡΠΎΠ±Π΅ΠΉ Ρ Π½Π΅ΡΠΎΠ³Π»Π°ΡΠΎΠ²Π°Π½Π½ΠΎΠΉ Π°Π»Π»ΠΎΠ·ΠΈΠΌΠ½ΠΎΠΉ ΠΈΠ·ΠΌΠ΅Π½ΡΠΈΠ²ΠΎΡΡΡΡ ΠΈ ΡΠ°Π·Π½Π°Ρ ΡΠΊΠ°Π½Π΅ΡΠΏΠ΅ΡΠΈΡΠΈΡΠ½ΠΎΡΡΡ ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΡΡΠΈΡ ΠΈΠ·ΠΎΡΠ΅ΡΠΌΠ΅Π½ΡΠΎΠ² — ΠΌΠΎΠΆΠ΅Ρ Π±ΡΡΡ ΠΎΠ±ΡΡΡΠ½Π΅Π½ΠΎ ΡΡΠ°ΡΡΠΈΠ΅ΠΌ Π°Π»ΡΡΠ΅ΡΠ½Π°ΡΠΈΠ²Π½ΠΎΠ³ΠΎ ΡΠΏΠ»Π°ΠΉΡΠΈΠ½Π³Π° Π² ΡΠΊΡΠΏΡΠ΅ΡΡΠΈΠΈ ΠΎΠ΄Π½ΠΎΠ³ΠΎ Π³Π΅Π½Π° ΠΠ€Π.
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