Π ΠΎΠ»Ρ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π±ΠΈΠΎΠ³Π΅Π½Π΅Π·Π΅ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ²
ΠΠ»Ρ Π±Π΅ΡΠΏΠΎΠ·Π²ΠΎΠ½ΠΎΡΠ½ΡΡ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΠΏΡΠΎΠ²Π΅Π΄Π΅Π½Π½ΡΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΠΏΠΎΠ·Π²ΠΎΠ»ΠΈΠ»ΠΈ Π²ΡΡΡΠ½ΠΈΡΡ ΡΠ½ΠΈΠ²Π΅ΡΡΠ°Π»ΡΠ½ΠΎΡΡΡ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠ° «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΡ Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΠΏΡΠ΅Π΄ΡΡΠ°Π²ΠΈΡΠ΅Π»Π΅ΠΉ Protostomia ΠΈ Deutorostomia. ΠΠ° ΠΏΡΠΈΠΌΠ΅ΡΠ΅ ΠΊΠ°ΠΌΠ±Π°Π»Ρ ΠΈ ΠΌΡΡΠΈ ΡΠ°ΡΡΠΌΠΎΡΡΠ΅Π½Π° Π²Π΅ΡΠΎΡΡΠ½ΠΎΡΡΡ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΡ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΠΏΠΎΠ·Π²ΠΎΠ½ΠΎΡΠ½ΡΡ . Π£ΡΠΈΡΡΠ²Π°Ρ, ΡΡΠΎ Ρ G. v errucosa «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΊΡΡ ΠΎΠ±Π½Π°ΡΡΠΆΠ΅Π½Ρ… Π§ΠΈΡΠ°ΡΡ Π΅ΡΡ >
- Π‘ΠΎΠ΄Π΅ΡΠΆΠ°Π½ΠΈΠ΅
- ΠΡΠ΄Π΅ΡΠΆΠΊΠ°
- ΠΠΈΡΠ΅ΡΠ°ΡΡΡΠ°
- ΠΡΡΠ³ΠΈΠ΅ ΡΠ°Π±ΠΎΡΡ
- ΠΠΎΠΌΠΎΡΡ Π² Π½Π°ΠΏΠΈΡΠ°Π½ΠΈΠΈ
Π‘ΠΎΠ΄Π΅ΡΠΆΠ°Π½ΠΈΠ΅
- 1. ΠΠΠΠΠ ΠΠΠ’ΠΠ ΠΠ’Π£Π Π«
- 1. 1. ΠΡΡΠΎΡΠΈΡ ΠΎΡΠΊΡΡΡΠΈΡ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΠΎΠΉ ΠΏΠ»Π°Π·ΠΌΡ
- 1. 2. ΠΠΊΡΠΏΠ΅ΡΠΈΠΌΠ΅Π½ΡΠ°Π»ΡΠ½ΡΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ
- 1. 3. Π‘ΡΡΡΠΊΡΡΡΠ½ΠΎ-ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΡΠ΅ Π°ΡΠΏΠ΅ΠΊΡΡ
- 1. 4. ΠΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΈ ΠΈ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΡΠ΅ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΡ
- 2. ΠΠΠ’ΠΠ ΠΠΠΠ« Π ΠΠΠ’ΠΠΠ«.&bdquo
- 2. 1. ΠΠ±ΡΠ΅ΠΊΡΡ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ
- 2. 1. 1. Dugesia tigrina
- 2. 1. 2. Ototyphlonemertes martynov
- 2. 1. 3. Strongylocentroyus nudus
- 2. 1. 4. Apostichopus japonicus
- 2. 1. 5. Pleuronectes asper
- 2. 1. 6. Mus musculus
- 2. 1. 7. Gracilaria verrucosa
- 2. 2. ΠΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎ-ΠΌΠΈΠΊΡΠΎΡΠΊΠΎΠΏΠΈΡΠ΅ΡΠΊΠΎΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅
- 2. 3. ΠΠΈΡΡΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ
- 2. 4. ΠΠΎΡΡΠΎΠΌΠ΅ΡΡΠΈΡ
- 2. 4. 2. 1. Π‘ΠΏΠ΅ΡΠΌΠ°ΡΠΎΠ³Π΅Π½Π½ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ A. japonicus
- 2. 4. 2. 2. ΠΠΎΠ³Π΅Π½Π½ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ A. japonicus
- 2. 4. 2. 3. ΠΠΌΠ±ΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ A. japonicus
- 2. 1. ΠΠ±ΡΠ΅ΠΊΡΡ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ
- 3. 1. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ
Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΈ ΠΌΠΈΡΠΎΡ
ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π³ΠΎΠ½ΠΈΠ°Π»ΡΠ½ΡΡ
ΠΊΠ»Π΅ΡΠΊΠ°Ρ
Π½Π΅ΠΊΠΎΡΠΎΡΡΡ
ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΠΎΡΠΎΡΡΡ
ΠΈ Π²ΡΠΎΡΠΈΡΠ½ΠΎΡΠΎΡΡΡ
- 3. 1. 1. Dugesia tigrina
- 3. 1. 2. Ototyplonemetes martynov
- 3. 1. 3. Strongylocentrotus nudus
- 3. 1. 4. Apostichopus japonicus
- 3. 1. 5. Pleuronectes asper
- 3. 1. 6. Mus musculus
- 3. 2. ΠΠΎΡΡΠΎΡΡΠ½ΠΊΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΠ΅ ΠΎΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠΈ ΠΏΠΎΠ»ΠΎΠ²ΡΡ
Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² Π² ΠΆΠΈΠ·Π½Π΅Π½Π½ΠΎΠΌ ΡΠΈΠΊΠ»Π΅ A. japonicus
- 3. 2. 1. ΠΠΎΠ³Π΅Π½Π΅Π·
- 3. 2. 2. Π‘ΠΏΠ΅ΡΠΌΠ°ΡΠΎΠ³Π΅Π½Π΅Π·
- 3. 2. 3. ΠΠΌΠ±ΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ
- 3. 3. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ
Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΈ ΠΌΠΈΡΠΎΡ
ΠΎΠ½Π΄ΡΠΈΠΉ Ρ ΠΌΠ°ΠΊΡΠΎΠ²ΠΎΠ΄ΠΎΡΠΎΡΠ»ΠΈ Gracilaria verrucosa
- 3. 3. 1. ΠΠ°ΠΌΠ΅ΡΠΎΠ³Π΅Π½Π΅Π·
- 3. 3. 2. ΠΠ°ΡΠΏΠΎΡΠΏΠΎΡΠΎΠ³Π΅Π½Π΅Π·
- 3. 3. 3. Π’Π΅ΡΡΠ°ΡΠΏΠΎΡΠΎΠ³Π΅Π½Π΅Π·
- 4. 1. Π€ΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΠ΅ Π³Π΅ΡΠΌΠΈΠ½Π°ΡΠΈΠ²Π½ΡΡ Π³ΡΠ°Π½ΡΠ» Π² ΠΏΠΎΠ·Π΄Π½ΠΈΡ ΠΎΠΎΡΠΈΡΠ°Ρ
- 4. 2. Π‘ΡΡΡΠΊΡΡΡΠ½ΠΎΠ΅ ΡΠΎΡΡΠΎΡΠ½ΠΈΠ΅ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² Π² ΡΠΌΠ±ΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ
- 4. 3. ΠΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠΈ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΡΠ΅ΡΠΌΠΈΠ½Π°Π»ΡΠ½ΡΡ Π³ΡΠ°Π½ΡΠ» ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ
- 4. 4. ΠΠΎΠΊΠ°Π»ΠΈΠ·Π°ΡΠΈΡ «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ Π² ΠΆΠΈΠ·Π½Π΅Π½Π½ΠΎΠΌ ΡΠΈΠΊΠ»Π΅ ΠΌΠ°ΠΊΡΠΎΠ²ΠΎΠ΄ΠΎΡΠΎΡΠ»Π΅ΠΉ
- 4. 5. ΠΡΠ΅ΠΌΠ΅ΠΉΠΎΡΠΈΡΠ΅ΡΠΊΠ°Ρ ΡΠΎΠ»Ρ ΡΠ²Π»Π΅Π½ΠΈΡ «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΡ
Π ΠΎΠ»Ρ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π±ΠΈΠΎΠ³Π΅Π½Π΅Π·Π΅ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² (ΡΠ΅ΡΠ΅ΡΠ°Ρ, ΠΊΡΡΡΠΎΠ²Π°Ρ, Π΄ΠΈΠΏΠ»ΠΎΠΌ, ΠΊΠΎΠ½ΡΡΠΎΠ»ΡΠ½Π°Ρ)
ΠΠΎΠ·Π½ΠΈΠΊΠ½ΠΎΠ²Π΅Π½ΠΈΠ΅ ΠΈ ΠΏΡΠΈΡΠΎΠ΄Π° ΠΊΠ»Π΅ΡΠΎΠΊ ΠΏΠΎΠ»ΠΎΠ²ΠΎΠΉ Π»ΠΈΠ½ΠΈΠΈ ΠΈΠ·ΡΡΠ°Π΅ΡΡΡ Π±ΠΎΠ»Π΅Π΅ 100 Π»Π΅Ρ. ΠΠΈΡΡΠ΅ΡΠ΅Π½ΡΠΈΠ°ΡΠΈΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΊΠ»Π΅ΡΠΎΠΊ Π·Π°Π²ΠΈΡΠΈΡ ΠΎΡ Π°ΠΊΡΠΈΠ²Π½ΠΎΡΡΠΈ ΡΠΈΡΠΎΠΏΠ»Π°Π·ΠΌΠ°ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΡΡΠ±ΡΡΠ°Π½ΡΠΈΠΈ, ΠΊΠΎΡΠΎΡΠ°Ρ ΠΏΠ΅ΡΠ΅Π΄Π°Π΅ΡΡΡ ΠΎΡ ΠΎΠ΄Π½ΠΎΠ³ΠΎ ΠΏΠΎΠΊΠΎΠ»Π΅Π½ΠΈΡ ΠΊ Π΄ΡΡΠ³ΠΎΠΌΡ, ΡΡΠΎ ΠΎΡΠ½ΠΎΠ²Π°Π½ΠΎ Π½Π° ΠΎΠ³ΡΠΎΠΌΠ½ΠΎΠΌ ΠΊΠΎΠ»ΠΈΡΠ΅ΡΡΠ²Π΅ Π½Π°Π±Π»ΡΠ΄Π΅Π½ΠΈΠΉ ΠΎΠΏΡΠ΅Π΄Π΅Π»Π΅Π½Π½ΡΡ ΡΠΈΡΠΎΠΏΠ»Π°Π·ΠΌΠ°ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΡΡΡΡΠΊΡΡΡ Π² ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ . ΠΠ° ΡΠ΅Π³ΠΎΠ΄Π½ΡΡΠ½ΠΈΠΉ Π΄Π΅Π½Ρ Π² ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ Π±ΠΎΠ»Π΅Π΅ ΡΠ΅ΠΌ Π²ΠΎΡΡΠΌΠΈΠ΄Π΅ΡΡΡΠΈ ΠΊΠ»Π°ΡΡΠΎΠ² ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΠΎΠ±Π½Π°ΡΡΠΆΠ΅Π½Ρ Ρ Π°ΡΠ°ΠΊΡΠ΅ΡΠ½ΡΠ΅ ΠΌΠΎΡΡΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΡΡΡΡΠΊΡΡΡΡ, Π½Π°Π·ΡΠ²Π°Π΅ΠΌΡΠ΅ ΠΏΠΎΠ»ΠΎΠ²ΡΠΌΠΈ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠ°ΠΌΠΈ. ΠΠΎΠ»ΠΎΠ²ΡΠ΅ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΡ Π²ΠΈΠ·ΡΠ°Π»ΠΈΠ·ΠΈΡΡΡΡΡΡ Π² ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎΠΌ ΠΌΠΈΠΊΡΠΎΡΠΊΠΎΠΏΠ΅ ΠΊΠ°ΠΊ ΡΠ΅ΠΌΠ½ΡΠ΅ ΡΠΈΠ±ΡΠΈΠ»Π»ΡΡΠ½ΠΎ-Π³ΡΠ°Π½ΡΠ»ΡΡΠ½ΡΠ΅ Π³ΡΠ°Π½ΡΠ»Ρ, Π½Π΅ ΠΎΠΊΡΡΠΆΠ΅Π½Π½ΡΠ΅ ΠΌΠ΅ΠΌΠ±ΡΠ°Π½ΠΎΠΉ, ΠΊΠΎΡΠΎΡΡΠ΅ Π²ΠΎ ΠΌΠ½ΠΎΠ³ΠΈΡ ΡΠ»ΡΡΠ°ΡΡ Π°ΡΡΠΎΡΠΈΠΈΡΠΎΠ²Π°Π½Ρ Ρ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΡΠΌΠΈ ΠΈ Π»ΠΎΠΊΠ°Π»ΠΈΠ·ΠΎΠ²Π°Π½Ρ Π² ΠΏΠ΅ΡΠΈΠ½ΡΠΊΠ»Π΅Π°ΡΠ½ΠΎΠΌ ΠΏΡΠΎΡΡΡΠ°Π½ΡΡΠ²Π΅. ΠΡΠΈ Π³ΡΠ°Π½ΡΠ»Ρ ΠΎΠ±Π½Π°ΡΡΠΆΠΈΠ²Π°ΡΡ Π² ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ Π½Π° Π²ΡΠ΅Ρ ΡΡΠ°Π΄ΠΈΡΡ ΡΠ°Π·Π²ΠΈΡΠΈΡ ΠΎΡ ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΡΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΊΠ»Π΅ΡΠΎΠΊ Π² ΡΠΌΠ±ΡΠΈΠΎΠ½Π΅ ΠΈ Π΄ΠΎ Π³Π°ΠΌΠ΅Ρ Π²ΠΎ Π²Π·ΡΠΎΡΠ»ΠΎΠΌ ΠΎΡΠ³Π°Π½ΠΈΠ·ΠΌΠ΅. ΠΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ Ρ ΠΌΠ½ΠΎΠ³ΠΈΡ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΡΠ°ΠΊ Π½Π°Π·ΡΠ²Π°Π΅ΠΌΡΠ΅ «nuage», Ρ ΡΠΎΠΌΠ°ΡΠΎΠΈΠ΄Π½ΡΠ΅ ΠΈ ΠΏΠ΅ΡΠΈΠ½ΡΠΊΠ»Π΅Π°ΡΠ½ΡΠ΅ ΡΠ΅Π»Π°, ΠΏΠΎΠ»ΡΡΠ½Π°Ρ ΠΏΠ»Π°Π·ΠΌΠ° ΠΈ ΠΈΠ½ΡΠ΅ΡΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΠΉ ΡΠ΅ΠΌΠ΅Π½Ρ ΡΠΎΠ΄Π΅ΡΠΆΠ°Ρ Π ΠΠ ΠΈ Π±Π΅Π»ΠΊΠΈ. ΠΠ΅ΡΠΌΠΎΡΡΡ Π½Π° ΡΠΎ, ΡΡΠΎ ΡΡΡΡΠΊΡΡΡΠ° ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΈΠ½ΡΠ΅Π½ΡΠΈΠ²Π½ΠΎ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π»Π°ΡΡ Ρ ΠΏΠΎΠΌΠΎΡΡΡ ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎ-ΠΌΠΈΠΊΡΠΎΡΠΊΠΎΠΏΠΈΡΠ΅ΡΠΊΠΈΡ ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΡΡ ΠΌΠ΅ΡΠΎΠ΄ΠΎΠ² Π² ΡΠ΅ΡΠ΅Π½ΠΈΠΈ 20 Π²Π΅ΠΊΠ°, ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌ ΠΈΡ ΡΡΠ½ΠΊΡΠΈΠΎΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π΄ΠΎ ΡΠΈΡ ΠΏΠΎΡ Π½Π΅ ΡΠ°ΡΡΠΈΡΡΠΎΠ²Π°Π½. ΠΠ³ΡΠΎΠΌΠ½ΠΎΠ΅ ΠΊΠΎΠ»ΠΈΡΠ΅ΡΡΠ²ΠΎ Π½Π°ΠΊΠΎΠΏΠ»Π΅Π½Π½ΡΡ Π΄Π°Π½Π½ΡΡ Π½Π°ΠΏΠΎΠ»Π½ΡΡΡ ΡΡΡΠ°Π½ΠΈΡΡ Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΠ½ΡΡ ΠΎΠ±Π·ΠΎΡΠΎΠ² Π²ΡΠ΅ Π±ΠΎΠ»ΡΡΠΈΠΌ ΠΊΠΎΠ»ΠΈΡΠ΅ΡΡΠ²ΠΎΠΌ ΠΏΠΎΠ΄ΡΠΎΠ±Π½ΠΎΡΡΠ΅ΠΉ, ΠΊΠΎΡΠΎΡΡΠ΅, ΡΠ΅ΠΌ Π½Π΅ ΠΌΠ΅Π½Π΅Π΅, ΠΏΠΎΠΊΠ° Π½Π΅ ΠΏΠΎΠ·Π²ΠΎΠ»ΡΡΡ ΠΏΠΎΡΡΡΠΎΠΈΡΡ ΡΡΡΠΎΠΉΠ½ΡΡ ΠΈ ΠΏΠΎΠ½ΡΡΠ½ΡΡ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΡ ΡΡΠ½ΠΊΡΠΈΠΎΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΈ Π±ΠΈΠΎΠ³Π΅Π½Π΅Π·Π° ΡΡΡΡΠΊΡΡΡ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΠΎΠΉ ΠΏΠ»Π°Π·ΠΌΡ.
ΠΠ±Π½Π°Π΄Π΅ΠΆΠΈΠ²Π°ΡΡΠΈΠΌ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½ΠΈΠ΅ΠΌ, ΡΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π²ΡΠΈΠΌΡΡ Π² 90-Ρ Π³ΠΎΠ΄Π°Ρ 20 Π²Π΅ΠΊΠ°, ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΎΡΠΊΡΡΡΠΈΠ΅ ΠΏΡΠΈΡΡΡΡΡΠ²ΠΈΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π ΠΠ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² ΡΠΎΡΡΠ°Π²Π΅ ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ², ΠΈΠΌΠ΅ΡΡΠΈΡ ΠΊΠ°ΠΊ ΠΊΠΎΠΌΠΏΠ°ΠΊΡΠ½ΡΡ, ΡΠ°ΠΊ ΠΈ Π΄ΠΈΡΠΏΠ΅ΡΠ³ΠΈΡΠΎΠ²Π°Π½Π½ΡΡ ΡΡΡΡΠΊΡΡΡΡ. Π ΠΎΠ»Ρ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠ΅ ΡΡΠ½ΠΊΡΠΈΠΎΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΠΎΠΉ ΠΏΠ»Π°Π·ΠΌΡ Π°ΠΊΡΠΈΠ²Π½ΠΎ Π΄ΠΈΡΠΊΡΡΠΈΡΡΠ΅ΡΡΡ Π² Π½Π°ΡΡΠ½ΠΎΠΉ ΠΏΠ΅ΡΠΈΠΎΠ΄ΠΈΠΊΠ΅. ΠΠ΄Π½ΠΈΠΌ ΠΈΠ· Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΡ Π²ΠΎΠΏΡΠΎΡΠΎΠ² ΡΠ²Π»ΡΠ΅ΡΡΡ ΡΠΎ, ΠΊΠ°ΠΊ ΠΌΠΎΠ»Π΅ΠΊΡΠ»Ρ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΡ Π½ΡΠΊΠ»Π΅ΠΈΠ½ΠΎΠ²ΡΡ ΠΊΠΈΡΠ»ΠΎΡ ΠΏΠΎΠΏΠ°Π΄Π°ΡΡ Π² ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π» ΠΏΠΎΠ»ΠΎΠ²ΠΎΠΉ ΡΡΠ±ΡΡΠ°Π½ΡΠΈΠΈ. Π‘ΠΎΠ³Π»Π°ΡΠ½ΠΎ Π½Π΅Π΄Π°Π²Π½Π΅ΠΉ Π½Π°Ρ ΠΎΠ΄ΠΊΠ΅ Π Π΅ΡΠ½ΠΎΠ²Π° ΠΈ ΡΠΎΠ°Π²Ρ. (Reunov et al., 2000) Ρ ΠΌΠΎΡΡΠΊΠΎΠ³ΠΎ Π΅ΠΆΠ° Antocidaris crassispina Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ Π½Π΅ΠΊΠΎΡΠΎΡΡΠ΅ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΈ ΡΠΏΠΎΡΠΎΠ±Π½Ρ ΡΡΡΠ°ΡΠΈΠ²Π°ΡΡ Π½Π°ΡΡΠΆΠ½ΡΡ ΠΌΠ΅ΠΌΠ±ΡΠ°Π½Ρ, Π² ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠ΅ ΡΠ΅Π³ΠΎ ΠΈΡ ΠΌΠ°ΡΡΠΈΠΊΡ ΠΏΠΎΠΏΠ°Π΄Π°Π΅Ρ Π² ΡΠΈΡΠΎΠΏΠ»Π°Π·ΠΌΡ. ΠΠ΅ ΠΈΡΠΊΠ»ΡΡΠ΅Π½ΠΎ, ΡΡΠΎ Π΄Π°Π½Π½ΡΠΉ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌ ΡΠ²Π»ΡΠ΅ΡΡΡ ΡΠ½ΠΈΠ²Π΅ΡΡΠ°Π»ΡΠ½ΡΠΌ ΡΠΏΠΎΡΠΎΠ±ΠΎΠΌ ΠΈΠ½ΡΡΠΎΠ΄ΡΡΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π² ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΏΠ»Π°Π·ΠΌΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ ΠΊΠ»Π΅ΡΠΎΠΊ Π½Π° ΡΡΠ°Π΄ΠΈΠΈ Π³ΠΎΠ½ΠΈΠ΅Π², ΠΈ ΡΡΠΎΡ Π²ΡΠ²ΠΎΠ΄ Π² ΡΠ»ΡΡΠ°Π΅ Π΅Π³ΠΎ ΠΏΡΠ°Π²ΠΎΠΌΠΎΡΠ½ΠΎΡΡΠΈ ΠΈΠ³ΡΠ°Π΅Ρ Π±ΠΎΠ»ΡΡΡΡ ΡΠΎΠ»Ρ Π² ΠΏΠΎΠ½ΠΈΠΌΠ°Π½ΠΈΠΈ ΡΠΏΠΎΡΠΎΠ±Π° «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ. ΠΠ΄Π½Π°ΠΊΠΎ ΡΠΎΠ»ΠΎΠΊΠ°Π»ΠΈΠ·ΠΎΠ²Π°Π½Π½ΠΎΡΡΡ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ ΠΈ ΡΡΡΡΠΊΡΡΡΠ½ΡΡ ΡΠ»Π΅ΠΌΠ΅Π½ΡΠΎΠ² ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΎΡΠΌΠ΅ΡΠ°ΡΡ Π² ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΠΏΠΎΠ»ΠΎΠ²ΠΎΠΉ Π»ΠΈΠ½ΠΈΠΈ ΠΈ Π½Π° Π΄ΡΡΠ³ΠΈΡ ΡΡΠ°ΠΏΠ°Ρ ΠΎΠ½ΡΠΎΠ³Π΅Π½Π΅Π·Π° ΠΈ ΡΡΠΎΠΌΡ ΡΠ°ΠΊΡΡ ΠΏΠΎΠΊΠ° Π½Π΅ Π½Π°Ρ ΠΎΠ΄ΡΡ ΠΎΠ±ΡΡΡΠ½Π΅Π½ΠΈΡ. Π’Π°ΠΊΠΈΠΌ ΠΎΠ±ΡΠ°Π·ΠΎΠΌ, Π²ΡΡΡΠ½Π΅Π½ΠΈΠ΅ ΡΠΎΠ»ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΎΠ΄Π½ΠΎΠΉ ΠΈΠ· Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΡ Π·Π°Π΄Π°Ρ, Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΡ Π½Π° Π²ΡΡΡΠ½Π΅Π½ΠΈΠ΅ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠ° ΡΡΠ½ΠΊΡΠΈΠΎΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² Π½Π° ΡΠ°Π·Π»ΠΈΡΠ½ΡΡ ΡΡΠ°Π΄ΠΈΡΡ ΠΆΠΈΠ·Π½Π΅Π½Π½ΠΎΠ³ΠΎ ΡΠΈΠΊΠ»Π° Ρ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΠΈ ΡΠ°ΡΡΠ΅Π½ΠΈΠΉ. ΠΡΠ΅Π²ΠΈΠ΄Π½ΠΎ, ΡΡΠΎ Π΄Π°Π»ΡΠ½Π΅ΠΉΡΠΈΠ΅ ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎ-ΠΌΠΈΠΊΡΠΎΡΠΊΠΎΠΏΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΠΎΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠ΅ΠΉ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Ρ ΠΏΠΎΠ»ΠΎΠ²ΡΠΌΠΈ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠ°ΠΌΠΈ Ρ ΡΠ°Π·Π»ΠΈΡΠ½ΡΡ ΠΏΡΠ΅Π΄ΡΡΠ°Π²ΠΈΡΠ΅Π»Π΅ΠΉ ΠΌΠ½ΠΎΠ³ΠΎΠΊΠ»Π΅ΡΠΎΡΠ½ΡΡ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΡΠ²Π»ΡΡΡΡΡ Π°ΠΊΡΡΠ°Π»ΡΠ½ΡΠΌΠΈ Π΄Π»Ρ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΌΠΎΡΡΠΎΡΡΠ½ΠΊΡΠΈΠΎΠ½Π°Π»ΡΠ½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΠΏΡΠΈΠΈ ΡΠΎΠ»ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π±ΠΈΠΎΠ³Π΅Π½Π΅Π·Π΅ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ².
Π¦Π΅Π»Ρ Π΄Π°Π½Π½ΠΎΠΉ ΡΠ°Π±ΠΎΡΡ ΡΠΎΡΡΠΎΠΈΡ Π² ΡΠ»ΡΡΡΠ°ΡΡΡΡΠΊΡΡΡΠ½ΠΎΠΌ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠΈ ΠΎΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠ΅ΠΉ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ Π½Π΅ΡΠΊΠΎΠ»ΡΠΊΠΈΡ ΠΏΡΠ΅Π΄ΡΡΠ°Π²ΠΈΡΠ΅Π»Π΅ΠΉ ΠΌΠ½ΠΎΠ³ΠΎΠΊΠ»Π΅ΡΠΎΡΠ½ΡΡ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ , ΡΠ°ΠΊΠΈΡ ΠΊΠ°ΠΊ ΠΏΠ»Π°Π½Π°ΡΠΈΡ Dugesia tigrina, Π½Π΅ΠΌΠ΅ΡΡΠΈΠ½Π° Ototyphlonemertes martynovi, ΠΌΠΎΡΡΠΊΠΎΠΉ Π΅ΠΆ Strongylocentrotus nudus, Π³ΠΎΠ»ΠΎΡΡΡΠΈΡ Apostichopus japonicus, ΠΏΡΠ΅Π΄ΡΡΠ°Π²ΠΈΡΠ΅Π»Ρ ΠΊΠ°ΠΌΠ±Π°Π»ΠΎΠ²ΡΡ ΡΡΠ± Pleuronectes asper ΠΈ ΠΌΡΡΡ Mus musculus.
ΠΠ»Ρ Π±Π΅ΡΠΏΠΎΠ·Π²ΠΎΠ½ΠΎΡΠ½ΡΡ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΠΏΡΠΎΠ²Π΅Π΄Π΅Π½Π½ΡΠ΅ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΠΏΠΎΠ·Π²ΠΎΠ»ΠΈΠ»ΠΈ Π²ΡΡΡΠ½ΠΈΡΡ ΡΠ½ΠΈΠ²Π΅ΡΡΠ°Π»ΡΠ½ΠΎΡΡΡ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠ° «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ³ΠΎ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΡ Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΠΏΡΠ΅Π΄ΡΡΠ°Π²ΠΈΡΠ΅Π»Π΅ΠΉ Protostomia ΠΈ Deutorostomia. ΠΠ° ΠΏΡΠΈΠΌΠ΅ΡΠ΅ ΠΊΠ°ΠΌΠ±Π°Π»Ρ ΠΈ ΠΌΡΡΠΈ ΡΠ°ΡΡΠΌΠΎΡΡΠ΅Π½Π° Π²Π΅ΡΠΎΡΡΠ½ΠΎΡΡΡ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΡ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΠΏΠΎΠ·Π²ΠΎΠ½ΠΎΡΠ½ΡΡ .
ΠΠ° ΠΏΡΠΈΠΌΠ΅ΡΠ΅ Π³ΠΎΠ»ΠΎΡΡΡΠΈΠΈ A. japonicus ΠΏΡΠ΅Π΄ΡΡΠΎΡΠ»ΠΎ Π²ΡΡΡΠ½ΠΈΡΡ, ΡΠ²Π»ΡΡΡΡΡ Π»ΠΈ Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ Π΅Π΄ΠΈΠ½ΡΡΠ²Π΅Π½Π½ΠΎΠΉ ΡΡΠ°Π΄ΠΈΠ΅ΠΉ Π² ΠΆΠΈΠ·Π½Π΅Π½Π½ΠΎΠΌ ΡΠΈΠΊΠ»Π΅, Π² ΠΊΠΎΡΠΎΡΠΎΠΌ ΠΏΡΠΎΠΈΡΡ ΠΎΠ΄ΠΈΡ «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ΅ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΠ΅ ΠΈΠ»ΠΈ ΠΆΠ΅ Π΄Π°Π½Π½ΠΎΠ΅ ΡΠ²Π»Π΅Π½ΠΈΠ΅ ΠΌΠΎΠΆΠ΅Ρ Π²ΡΡΡΠ΅ΡΠ°ΡΡΡΡ ΠΈ Π² Π΄ΡΡΠ³ΠΈΡ ΡΡΠ°Π΄ΠΈΡΡ , ΠΊΠΎΡΠΎΡΡΠΌΠΈ ΡΠ²Π»ΡΡΡΡΡ ΠΏΠΎΡΡΠ³ΠΎΠ½ΠΈΠ°Π»ΡΠ½ΡΠ΅ ΡΡΠ°ΠΏΡ Π³Π°ΠΌΠ΅ΡΠΎΠ³Π΅Π½Π΅Π·Π° ΠΈ ΡΠΌΠ±ΡΠΈΠΎΠ³Π΅Π½Π΅Π·.
ΠΠ° ΠΏΡΠΈΠΌΠ΅ΡΠ΅ ΠΊΡΠ°ΡΠ½ΠΎΠΉ Π²ΠΎΠ΄ΠΎΡΠΎΡΠ»ΠΈ Gracilaria verrucosa, ΠΆΠΈΠ·Π½Π΅Π½Π½ΡΠΉ ΡΠΈΠΊΠ» ΠΊΠΎΡΠΎΡΠΎΠΉ Π²ΠΊΠ»ΡΡΠ°Π΅Ρ ΠΌΠ΅ΠΉΠΎΡΠΈΡΠ΅ΡΠΊΡΡ ΡΠ°Π·Ρ ΡΠΏΠΎΡΠΎΠ³Π΅Π½Π΅Π·Π° ΠΈ ΡΠ΅ΡΠ΅Π΄ΡΠ΅ΡΡΡ Ρ Π½Π΅ΠΌΠ΅ΠΉΠΎΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΡΠ°Π·ΠΎΠΉ Π³Π°ΠΌΠ΅ΡΠΎΠ³Π΅Π½Π΅Π·Π°, Π½Π΅ΠΎΠ±Ρ ΠΎΠ΄ΠΈΠΌΠΎ Π±ΡΠ»ΠΎ Π²ΡΡΡΠ½ΠΈΡΡ, ΡΠ²Π»ΡΠ΅ΡΡΡ Π»ΠΈ «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ΅ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΠ΅ Ρ Π°ΡΠ°ΠΊΡΠ΅ΡΠ½ΡΠΌ ΡΠΎΠ»ΡΠΊΠΎ Π΄Π»Ρ ΠΌΠ΅ΠΉΠΎΡΠΈΡΠ΅ΡΠΊΠΈΡ ΠΊΠ»Π΅ΡΠΎΠΊ, ΠΈΠ»ΠΈ ΠΆΠ΅ Π΄Π°Π½Π½ΠΎΠ΅ ΡΠ²Π»Π΅Π½ΠΈΠ΅ Π²ΡΡΡΠ΅ΡΠ°Π΅ΡΡΡ ΠΈ Π² ΠΏΡΠΎΡΠ΅ΡΡΠ΅ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π³Π°ΠΌΠ΅Ρ, ΠΊΠΎΡΠΎΡΠΎΠ΅ Ρ Rhodophyta Π½Π΅ ΡΠΎΠΏΡΠΎΠ²ΠΎΠΆΠ΄Π°Π΅ΡΡΡ ΠΌΠ΅ΠΉΠΎΠ·ΠΎΠΌ.
ΠΠ»Ρ Π΄ΠΎΡΡΠΈΠΆΠ΅Π½ΠΈΡ Π΄Π°Π½Π½ΡΡ ΡΠ΅Π»Π΅ΠΉ ΠΏΡΠ΅Π΄ΠΏΠΎΠ»Π°Π³Π°Π»ΠΎΡΡ ΡΠ΅ΡΠΈΡΡ ΡΠ»Π΅Π΄ΡΡΡΠΈΠ΅ Π·Π°Π΄Π°ΡΠΈ:
1. Π‘ ΠΏΠΎΠΌΠΎΡΡΡ ΡΠ»ΡΡΡΠ°ΡΡΡΡΠΊΡΡΡΠ½ΠΎΠ³ΠΎ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΡΠ΅ΡΠΈΠΉΠ½ΡΡ ΡΡΠ΅Π·ΠΎΠ² ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΡ «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΊΡΡ Π² Π³ΠΎΠ½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΠΏΠ»Π°Π½Π°ΡΠΈΠΈ D. tigrina, Π½Π΅ΠΌΠ΅ΡΡΠΈΠ½Ρ Π. martynovi, ΠΌΠΎΡΡΠΊΠΎΠ³ΠΎ Π΅ΠΆΠ° S. nudus, Π³ΠΎΠ»ΠΎΡΡΡΠΈΠΈ A. japonicus, ΠΊΠ°ΠΌΠ±Π°Π»ΠΎΠ²ΠΎΠΉ ΡΡΠ±Ρ Π . asper ΠΈ ΠΌΡΡΠΈ Π. musculus.
2. ΠΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎ-ΠΌΠΈΠΊΡΠΎΡΠΊΠΎΠΏΠΈΡΠ΅ΡΠΊΠΈ ΠΈ ΠΌΠΎΡΡΠΎΠΌΠ΅ΡΡΠΈΡΠ΅ΡΠΊΠΈ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΡ ΠΎΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠΈ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² ΠΎΠΎΠ³Π΅Π½Π½ΡΡ , ΡΠΏΠ΅ΡΠΌΠ°ΡΠΎΠ³Π΅Π½Π½ΡΡ ΠΈ ΡΠΌΠ±ΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ Π³ΠΎΠ»ΠΎΡΡΡΠΈΠΈ Π. japonicus.
3. ΠΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎ-ΠΌΠΈΠΊΡΠΎΡΠΊΠΎΠΏΠΈΡΠ΅ΡΠΊΠΈ ΠΈ ΠΌΠΎΡΡΠΎΠΌΠ΅ΡΡΠΈΡΠ΅ΡΠΊΠΈ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΡ ΠΎΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠΈ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΡ ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² ΡΠΏΠΎΡΠΎΠ³Π΅Π½Π΅Π·Π΅ ΠΈ Π³Π°ΠΌΠ΅ΡΠΎΠ³Π΅Π½Π΅Π·Π΅ ΠΊΡΠ°ΡΠ½ΠΎΠΉ Π²ΠΎΠ΄ΠΎΡΠΎΡΠ»ΠΈ G. verrucosa.
Π Π΅ΡΠ΅Π½ΠΈΠ΅ Π΄Π°Π½Π½ΡΡ Π·Π°Π΄Π°Ρ ΡΠΏΠΎΡΠΎΠ±ΡΡΠ²ΡΠ΅Ρ Π²ΡΡΡΠ½Π΅Π½ΠΈΡ ΡΠΎΠ»ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ Π² Π±ΠΈΠΎΠ³Π΅Π½Π΅Π·Π΅ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΠΎΠΉ ΠΏΠ»Π°Π·ΠΌΡ.
1. ΠΠΠΠΠ ΠΠΠ’ΠΠ ΠΠ’Π£Π Π«.
ΠΠ«ΠΠΠΠ«.
1. ΠΠ°ΡΠ°Π»ΡΠ½ΡΠΌ ΡΡΠ°ΠΏΠΎΠΌ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π·Π°ΡΠΎΠ΄ΡΡΠ΅Π²ΠΎΠΉ ΠΏΠ»Π°Π·ΠΌΡ Π² ΠΆΠΈΠ·Π½Π΅Π½Π½ΠΎΠΌ ΡΠΈΠΊΠ»Π΅ ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΎΠ±ΡΠ°Π·ΠΎΠ²Π°Π½ΠΈΠ΅ Π³Π΅ΡΠΌΠΈΠ½Π°ΡΠΈΠ²Π½ΡΡ Π³ΡΠ°Π½ΡΠ» Π² ΠΏΠΎΠ·Π΄Π½Π΅ΠΌ ΠΎΠΎΠ³Π΅Π½Π΅Π·Π΅. Π£ A. japonicus Π³Π΅ΡΠΌΠΈΠ½Π°ΡΠΈΠ²Π½ΡΠ΅ Π³ΡΠ°Π½ΡΠ»Ρ ΡΠΎΡΠΌΠΈΡΡΡΡΡΡ ΠΈΠ· * ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ, ΠΊΠΎΠ½ΡΠ°ΠΊΡΠΈΡΡΡΡΠΈΡ Ρ ΡΡΠ±ΡΡΠ°Π½ΡΠΈΠ΅ΠΉ ΡΠ΅Π»Π° ΠΠ°Π»ΡΠ±ΠΈΠ°Π½ΠΈ.
2. Π ΡΠΌΠ±ΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ A. japonicus ΠΏΠΎΠ»ΠΎΠ²ΡΠ΅ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΡ, Π²ΠΎΠ·Π½ΠΈΠΊΡΠΈΠ΅ Π² ΠΎΠΎΡΠΈΡΠ°Ρ , ΠΏΡΠΈΡΡΡΡΡΠ²ΡΡΡ Π² ΠΈΠ½ΡΠ°ΠΊΡΠ½ΠΎΠΉ ΡΠΎΡΠΌΠ΅ ΠΈ Π½Π΅ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΡΡΡ Ρ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΡΠΌΠΈ.
3. Π Π³ΠΎΠ½ΠΈΠ°Π»ΡΠ½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ A. japonicus, Π° ΡΠ°ΠΊΠΆΠ΅ ΡΠ°ΠΊΠΈΡ ΠΏΡΠ΅Π΄ΡΡΠ°Π²ΠΈΡΠ΅Π»Π΅ΠΉ ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΠΎΡΠΎΡΡΡ ΠΈ Π²ΡΠΎΡΠΈΡΠ½ΠΎΡΠΎΡΡΡ , ΠΊΠ°ΠΊ D. tigrina, Π. martynovi, S. nudus, P. asper ΠΈ M. musculus, ΠΏΡΠΎΠΈΡΡ ΠΎΠ΄ΠΈΡ ΡΡΠ°Π½ΡΡΠΎΡΠΌΠ°ΡΠΈΡ ΠΊΠΎΠΌΠΏΠ°ΠΊΡΠ½ΡΡ Π³Π΅ΡΠΌΠΈΠ½Π°ΡΠΈΠ²Π½ΡΡ Π³ΡΠ°Π½ΡΠ» Π² ΡΠ°ΡΡΠ΅ΡΠ½Π½ΠΎΠ΅ ΡΠΎΡΡΠΎΡΠ½ΠΈΠ΅. ΠΠ·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΡΡ, Ρ Π΄Π°Π½Π½ΠΎΠΉ ΡΡΠ±ΡΡΠ°Π½ΡΠΈΠ΅ΠΉ («nuage») ΡΠ°ΡΡΡ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠΉ ΠΊΠ»Π΅ΡΠΊΠΈ ΡΠΊΡΠΊΡΠ΅ΡΠΈΡΡΠ΅Ρ ΡΠΎΠ΄Π΅ΡΠΆΠ°ΡΠΈΠΉ ΠΊΡΠΈΡΡΡ ΠΌΠ°ΡΡΠΈΠΊΡ.
4. Π£ΡΠΈΡΡΠ²Π°Ρ, ΡΡΠΎ ΡΠ°ΠΊΠΈΠ΅ ΡΡΡΡΠΊΡΡΡΡ ΠΊΠ°ΠΊ Π³Π΅ΡΠΌΠΈΠ½Π°ΡΠΈΠ²Π½ΡΠ΅ Π³ΡΠ°Π½ΡΠ»Ρ, «nuage», «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΊΡΡ ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΏΡΠΎΠΈΠ·Π²ΠΎΠ΄Π½ΡΠ΅, Π½Π΅ Π²ΡΡΡΠ΅ΡΠ°ΡΡΡΡ Π² ΠΏΠΎΡΠ»Π΅Π΄ΡΡΡΠΈΡ ΡΡΠ°Π΄ΠΈΡΡ Π³Π°ΠΌΠ΅ΡΠΎΠ³Π΅Π½Π΅Π·Π° Π. Japonicus, ΠΌΠΎΠΆΠ½ΠΎ ΡΡΠΈΡΠ°ΡΡ, ΡΡΠΎ Π³Π΅Π½ΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ ΡΠ²Π»ΡΡΡΡΡ Π΅Π΄ΠΈΠ½ΡΡΠ²Π΅Π½Π½ΡΠΌ ΡΡΠ°ΠΏΠΎΠΌ ΠΆΠΈΠ·Π½Π΅Π½Π½ΠΎΠ³ΠΎ ΡΠΈΠΊΠ»Π°, Π² ΠΊΠΎΡΠΎΡΠΎΠΌ ΠΏΡΠΎΠΈΡΡ ΠΎΠ΄ΠΈΡ ΡΠ°ΡΡΠ΅ΠΈΠ²Π°Π½ΠΈΠ΅ Π³Π΅ΡΠΌΠΈΠ½Π°ΡΠΈΠ²Π½ΡΡ Π³ΡΠ°Π½ΡΠ» ΠΈ ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΡ ΠΏΡΠΎΠΈΠ·Π²ΠΎΠ΄Π½ΡΡ .
5. ΠΠ°Π»ΠΈΡΠΈΠ΅ ΠΏΠΎΠ»ΠΎΠ²ΡΡ Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½Π°Π½ΡΠΎΠ² ΡΠ²ΠΎΠΉΡΡΠ²Π΅Π½Π½ΠΎ Π½Π΅ ΡΠΎΠ»ΡΠΊΠΎ ΠΌΠ½ΠΎΠ³ΠΎΠΊΠ»Π΅ΡΠΎΡΠ½ΡΠΌ ΠΆΠΈΠ²ΠΎΡΠ½ΡΠΌ, Π½ΠΎ ΠΈ ΠΌΠ°ΠΊΡΠΎΠ²ΠΎΠ΄ΠΎΡΠΎΡΠ»ΠΈ G. verrucosa, ΡΡΠΎ ΡΠ²ΠΈΠ΄Π΅ΡΠ΅Π»ΡΡΡΠ²ΡΠ΅Ρ ΠΎ Π²ΠΎΠ·ΠΌΠΎΠΆΠ½ΠΎΠΉ ΡΠ½ΠΈΠ²Π΅ΡΡΠ°Π»ΡΠ½ΠΎΡΡΠΈ ΠΈ ΡΠ²ΠΎΠ»ΡΡΠΈΠΎΠ½Π½ΠΎΠΌ ΠΊΠΎΠ½ΡΠ΅ΡΠ²Π°ΡΠΈΠ·ΠΌΠ΅ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠ°, Π΄Π΅ΡΠ΅ΡΠΌΠΈΠ½ΠΈΡΡΡΡΠ΅Π³ΠΎ ΠΏΠΎΠ»ΠΎΠ²ΡΠ΅ ΠΊΠ»Π΅ΡΠΊΠΈ ΠΆΠΈΠ²ΠΎΡΠ½ΡΡ ΠΈ ΡΠ°ΡΡΠ΅Π½ΠΈΠΉ.
6. Π£ΡΠΈΡΡΠ²Π°Ρ, ΡΡΠΎ Ρ G. v errucosa «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΊΡΡ ΠΎΠ±Π½Π°ΡΡΠΆΠ΅Π½Ρ Π½Π΅ Π² Π³Π°ΠΌΠ΅ΡΠΎΠ³Π΅Π½Π½ΡΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ , Π° Π² ΠΏΡΠ΅ΠΌΠ΅ΠΉΠΎΡΠΈΡΠ΅ΡΠΊΠΈΡ ΠΊΠ»Π΅ΡΠΊΠ°Ρ ΡΠΏΠΎΡΠ°Π½Π³ΠΈΠ΅Π², ΠΌΠΎΠΆΠ½ΠΎ ΡΡΠΈΡΠ°ΡΡ, ΡΡΠΎ «nuage''-ΠΌΠΈΡΠΎΡ ΠΎΠ½Π΄ΡΠΈΠ°Π»ΡΠ½ΠΎΠ΅ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΠ΅ ΠΈΠΌΠ΅Π΅Ρ Π·Π½Π°ΡΠ΅Π½ΠΈΠ΅ Π½Π΅ ΡΡΠΎΠ»ΡΠΊΠΎ Π΄Π»Ρ ΡΠΎΡΠΌΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π³Π°ΠΌΠ΅Ρ, ΡΠΊΠΎΠ»ΡΠΊΠΎ Π΄Π»Ρ ΡΡΠΈΠΌΡΠ»ΡΡΠΈΠΈ ΠΏΡΠΎΡΠ΅ΡΡΠ° ΠΌΠ΅ΠΉΠΎΠ·Π°.
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